LECTURE 6: BIOSPHERE
A planet that can sustain life is termed habitable, even if life did not originate there. The Earth provides the (currently understood) requisite conditions of liquid water, an environment where complex organic molecules can assemble, and sufficient energy to sustain metabolism.[95] The distance of the Earth from the Sun, as well as its orbital eccentricity, rate of rotation, axial tilt, geological history, sustaining atmosphere and protective magnetic field all contribute to the conditions necessary to originate and sustain life on this planet.[96]
The biosphere is the part of the Earth, including air, land, surface rocks, and water, within which life occurs, and which biotic processes in turn alter or transform. From the broadest biophysiological point of view, the biosphere is the global ecological system integrating all living beings and their relationships, including their interaction with the elements of the lithosphere, hydrosphere, and atmosphere. This biosphere is postulated to have evolved, beginning through a process of biogenesis or biopoesis, at least some 3.5 billion years ago.
Biomass accounts for about 3.7 kg carbon per square meter of the earth's surface averaged over land and sea, making a total of about 1900 gigatonnes of carbon.
Some life scientists and earth scientists use biosphere in a more limited sense. For example, geochemists define the biosphere as being the total sum of living organisms (the "biomass" or "biota" as referred to by biologists and ecologists). In this sense, the biosphere is but one of four separate components of the geochemical model, the other three being lithosphere, hydrosphere, and atmosphere. The narrow meaning used by geochemists is one of the consequences of specialization in modern science. Some might prefer the word ecosphere, coined in the 1960s, as all encompassing of both biological and physical components of the planet.
The Second International Conference on Closed Life Systems defined biospherics as the science and technology of analogs and models of Earth's biosphere; i.e., artificial Earth-like biospheres. Others may include the creation of artificial non-Earth biospheres — for example, human-centered biospheres or a native Martian biosphere — in the field of biospherics.
The concept that the biosphere is itself a living organism, either actually or metaphorically, is known as the Gaia hypothesis.
James Lovelock, an atmospheric scientist from the United Kingdom, proposed the Gaia hypothesis to explain how biotic and abiotic factors interact in the biosphere. This hypothesis considers Earth itself a kind of living organism. Its atmosphere, geosphere, and hydrosphere are cooperating systems that yield a biosphere full of life. in the early 1970s, Lynn Margulis, a microbiologist from the United States, added to the hypothesis specifically noting the ties between the biosphere and other Earth systems. For example, when carbon dioxide levels increase in the atmosphere, plants grow more quickly. As their growth continue, they remove more and more carbon dioxide from the atmosphere.
Many scientists are now devoting their careers to organizing new fields of study, such as geobiology and geomicrobiology, to examine these intriguing relationships .
Nearly every part of the planet, from the polar ice caps to the Equator, supports life of some kind. Recent advances in microbiology have demonstrated that microbes live deep beneath the Earth's terrestrial surface, and that the total mass of microbial life in so-called "uninhabitable zones" may, in biomass, exceed all animal and plant life on the surface.
The actual thickness of the biosphere on earth is hard to measure. Birds typically fly at altitudes of 650 to 2000 meters, and fish that live deep underwater can be found down to -8,372 meters in the Puerto Rico Trench.
There are more extreme examples for life on the planet: Rüppell's Vulture has been found at altitudes of 11,300 meters; Bar-headed Geese migrate at altitudes of at least 8,300 meters (over Mount Everest); Yaks live at elevations between 3,200 to 5,400 meters above sea level; mountain goats live up to 3,050 meters. Herbivorous animals at these elevations depend on lichens, grasses, and herbs but the biggest tree is the Tine palm or mountain coconut found 3,400 meters above sea level.
Microscopic organisms live at such extremes that, taking them into consideration puts the thickness of the biosphere much greater, but at minimum it extends from 5,400 meters above sea level to at least 9,000 meters below sea level.
Our biosphere is divided into a number of biomes, inhabited by broadly similar flora and fauna. On land, biomes are separated primarily by latitude. Terrestrial biomes lying within the Arctic and Antarctic Circles are relatively barren of plant and animal life, while most of the more populous biomes lie near the equator. Terrestrial organisms in temperate and Arctic biomes have relatively small amounts of total biomass, smaller energy budgets, and display prominent adaptations to cold, including world-spanning migrations, social adaptations, homeothermy, estivation and multiple layers of insulation.
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