BIO 103

 

Paleozoic Life

 

 

 

 

 

Back

 

 

 

LECTURE 1: THE CALENDAR OF LIFE ON EARTH

 The basic timeline is a 4.6 billion year old Earth, with (very approximately):

 

4567.17 Ma The planet Earth forms from the accretion disc revolving around the young Sun.
4533 Ma The planet Earth and the planet Theia collide, sending countless moonlets into orbit around the young Earth. These moonlets eventually coalesce to form the Moon. The gravitational pull of the new Moon stabilises the Earth's fluctuating axis of rotation and sets up the conditions for the formation of life.[1]
4100 Ma The surface of the Earth cools enough for the crust to solidify. The atmosphere and the oceans form.[2]PAH infall, and Iron-Sulfide synthesis along deep ocean platelet boundaries, may have led to the RNA world of competing metabolising organic compounds.
4500 - 2500 Ma The earliest life appears, possibly derived from self-reproducing RNA molecules. The replication of these organisms requires resources like energy, space, and smaller building blocks, which soon become limited, resulting in competition. Natural selection favours those molecules which are more efficient at replication. DNA molecules then take over as the main replicators. They soon develop inside enclosing membranes which provide a stable physical and chemical environment conducive to their replication: proto-cells.
3900 Ma Late Heavy Bombardment: peak rate of impact events upon the inner planets by meteors. This constant disturbance probably obliterated any life that had already evolved, as the oceans boiled away completely; conversely, life may have been transported to Earth by a meteor. [3]
3900 - 2500 Ma Cells resembling prokaryotes appear. These first organisms are chemoautotrophs: they use carbon dioxide as a carbon source and oxidize inorganic materials to extract energy. Later, prokaryotes evolve glycolysis, a set of chemical reactions that free the energy of organic molecules such as glucose. Glycolysis generates ATP molecules as short-term energy currency, and ATP continue to be used in almost all organisms, unchanged, to this day.
3500 Ma Lifetime of the last universal ancestor; the split between the bacteria and the archaea occurs.

Bacteria develop primitive forms of photosynthesis which at first do not produce oxygen. These organisms generate ATP by exploiting a proton gradient, a mechanism still used in virtually all organisms.

3000 Ma Photosynthesizing cyanobacteria evolve; they use water as a reducing agent, thereby producing oxygen as waste product. The oxygen initially oxidizes dissolved iron in the oceans, creating iron ore. The oxygen concentration in the atmosphere subsequently rises, acting as a poison for many bacteria. The moon is still very close to the earth and causes tides 1000 feet high. The earth is continually wracked by hurricane force winds. These extreme mixing influences are thought to stimulate evolutionary processes. (See Oxygen Catastrophe)
530 Ma The first known footprints on land date to 530 Ma, indicating that early animal explorations may have predated the development of terrestrial plants.[5]
475 Ma The first primitive plants move onto land,[6][citation needed] having evolved from green algae living along the edges of lakes.[7] They are accompanied by fungi, which may have aided the colonisation of land through symbiosis.
363 Ma By the start of the Carboniferous period, the Earth begins to be recognisable. Insects roamed the land and would soon take to the skies; sharks predated the oceans,[8] and vegetation covered the land, with seed-bearing plants and forests soon to flourish.

Four-limbed tetrapods gradually gain adaptations which will help them occupy a terrestrial life-habit.

251.4Ma The Permian-Triassic extinction event eliminates over 95% of species. This "clearing of the slate" may have led to an ensuing diversification.

 

READING FOR THE NEXT LECTURE

 

Return to BIO 103

 

 

 

     Prof. Torgersen